Skip to main content
Submit
About
Editorial Board
PNAS Staff
FAQ
Accessibility Statement
Rights and Permissions
Site Map
Contact
Journal Club
Subscribe
Subscription Rates
Subscriptions FAQ
Open Access
Recommend PNAS to Your Librarian
Log in
My Cart
Main menu
Home
Articles
Current
Latest Articles
Special Features
Colloquia
Collected Articles
PNAS Classics
Archive
Front Matter
News
For the Press
Highlights from Latest Articles
PNAS in the News
Podcasts
Authors
Information for Authors
Purpose and Scope
Editorial and Journal Policies
Submission Procedures
For Reviewers
Author FAQ
Submit
About
Editorial Board
PNAS Staff
FAQ
Accessibility Statement
Rights and Permissions
Site Map
Contact
Journal Club
Subscribe
Subscription Rates
Subscriptions FAQ
Open Access
Recommend PNAS to Your Librarian
User menu
Log in
My Cart
Search
Search for this keyword
Search
Advanced search
Search for this keyword
Search
Advanced Search
Home
Articles
Current
Latest Articles
Special Features
Colloquia
Collected Articles
PNAS Classics
Archive
Front Matter
News
For the Press
Highlights from Latest Articles
PNAS in the News
Podcasts
Authors
Information for Authors
Purpose and Scope
Editorial and Journal Policies
Submission Procedures
For Reviewers
Author FAQ
New Research In
Physical Sciences
Featured Portals
Physics
Chemistry
Sustainability Science
Articles by Topic
Applied Mathematics
Applied Physical Sciences
Astronomy
Computer Sciences
Earth, Atmospheric, and Planetary Sciences
Engineering
Environmental Sciences
Mathematics
Statistics
Social Sciences
Featured Portals
Anthropology
Sustainability Science
Articles by Topic
Economic Sciences
Environmental Sciences
Political Sciences
Psychological and Cognitive Sciences
Social Sciences
Biological Sciences
Featured Portals
Sustainability Science
Articles by Topic
Agricultural Sciences
Anthropology
Applied Biological Sciences
Biochemistry
Biophysics and Computational Biology
Cell Biology
Developmental Biology
Ecology
Environmental Sciences
Evolution
Genetics
Immunology and Inflammation
Medical Sciences
Microbiology
Neuroscience
Pharmacology
Physiology
Plant Biology
Population Biology
Psychological and Cognitive Sciences
Sustainability Science
Systems Biology
Site Map
Articles and Research
Table of Contents
Latest Articles
Issues Archive
Alerts
RSS
Collected Articles
Front Matter magazine
PNAS Portals
Classics
Teaching Resources
Anthropology
Chemistry
Physics
Sustainability Science
About PNAS
PNAS Updates
About FAQ
Editorial Board
Staff Directory
For the Press
Rights and Permissions
Information Brochure
Journal Metrics
Developing Countries
Cozzarelli Prize
About Direct Submission
RSS
Reprints
Terms of Use
For Subscribers
Subscriber FAQ
Subscription Rates
Open Access
Order Forms
Subscription Order Form (PDF)
Order print-on-demand issues
For Authors
Author FAQ
Purpose and Scope
Editorial and Journal Policies
Competing Interest
Submission Procedures
Manuscript Format
For Reviewers
LaTeX
Production Process
Publication Fees
Licenses for PNAS Articles
Call for Papers
Information for Authors (PDF)
[09/19]